Two new genera and species of permineralized tree ferns, Natalipteris wildei and Kwazulupteris schaarschmidtii, are defined and described in detail. Natalipteris wildei is a solenostelic stem without well-developed sclerenchyma sheaths in their vascular strands, and has a single vascular strand in the petiole bases, which are somewhat similar to the one present in ...